 | Gray Wolf: Encyclopedia II - Gray Wolf - Social structure
Gray Wolf - Social structure
Gray Wolf - Packs
Wolves function as social predators and hunt in packs organized according to a strict social hierarchy and led by an alpha male and alpha female. This social structure was originally thought to allow the wolf to take prey many times its size. However, emerging new theories suggest the pack strategy has less to do with hunting than with reproductive success.
The size of the pack may change over time and is controlled by several factors, including habitat, personalities of individual wolves within a pack, and food supply. Packs can contain between two and 20 wolves, though an average pack consists of six or seven.[4][5] The hierarchy of the pack is relatively strict, with the alphas at the top and the omega at the bottom. The hierarchy affects all activity in the pack, from which wolf eats first to which is allowed to breed (generally only the alpha pair).
New packs are formed when a wolf leaves its birth pack and claims a territory. Lone wolves searching for other individuals can travel very long distances seeking out suitable territories. Dispersing individuals must avoid the territories of other wolves because intruders on occupied territories are chased away or killed, a behavior that may explain wolf "predation" of dogs. Most dogs, except perhaps large, specially bred attack dogs, do not stand much of a chance against a pack of wolves protecting its territory from an unwanted intrusion.
Gray Wolf - Rank order
The alpha pair has the greatest amount of social freedom among all the pack members, but they are not "leaders" in the human sense of the term. The alphas do not give the other wolves orders; rather, they simply have the most freedom in choosing where to go, what to do, and when to do it. The rest of the pack usually follows.
In larger packs, there may be also be a beta wolf or wolves – a "second-in-command" to the alphas. In addition, one wolf typically assumes the role of omega, the lowest-ranking member of a pack.[6] These individuals absorb the greatest amount of aggression from the rest of the pack, and consequently enjoy comparatively few individual privileges.
While most alpha pairs are monogamous with each other, there are exceptions.[7] An alpha animal may preferentially mate with a lower-ranking animal, especially if the other alpha is closely related (a brother or sister, for example). The death of one alpha does not affect the status of the other alpha, who will quickly take another mate. Usually, only the alpha pair is able to successfully rear a litter of pups (other wolves in a pack may breed, and may even produce pups, but usually they lack the freedom or the resources to raise the pups to maturity). All the wolves in the pack assist in raising wolf pups. Some mature individuals, usually females, may choose to stay in the original pack so as to reinforce it and help rear more pups. Most, males particularly, will disperse, however.
Rank order is established and maintained through a series of ritualized fights and posturing best described as ritual bluffing. Wolves prefer psychological warfare to physical confrontations, meaning that high-ranking status is based more on personality or attitude than on size or physical strength. Rank, who holds it, and how it is enforced varies widely between packs and between individual animals. In large packs full of easygoing wolves, or in a group of juvenile wolves, rank order may shift almost constantly, or even be circular (e.g., animal A dominates animal B, who dominates animal C, who dominates animal A).
Loss of rank can happen gradually or suddenly. An older wolf may simply choose to give way when an ambitious challenger presents itself, yielding its position without bloodshed. On the other hand, the challenged individual may choose to fight back, with varying degrees of intensity. While the majority of wolf aggression is non-damaging and ritualized, a high-stakes fight can easily result in injury for either or both parties. The loser of such a confrontation is frequently chased away from the pack or, rarely, may be killed as other aggressive wolves contribute to the insurgency. This kind of dominance encounter is more common in the winter months, when mating occurs.
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